Royal Society Awards & Honours

The Royal Society is a Fellowship of the world’s most eminent scientists and is the oldest scientific academy in continuous existence. Each year the Royal Society recognizes excellence in science and technology through its medals, awards and prize lectures and the election of new Fellows. Fellows are elected through a peer review process. The main criterion for election as a Fellow is scientific excellence. The following LMB scientists have been recognized by the Royal Society through election to the Fellowship and through various medals, awards and prize lectures. All the citations for awards, medals and prize lectures have been copied from the Royal Society website.


Fellows of the Royal Society
Andrew McKenzie 2017
John SutherlandJohn Sutherland 2017
Roger WilliamsRoger Williams 2017
Ramanujan HegdeRamanujan Hegde 2016
KJ Patel 2015
Sean Munro 2011
Michael Hastings 2010
Jan Löwe 2008
Harvey McMahon 2008
Brad Amos 2007
Daniela Rhodes 2007
David Barford 2006
Matthew Freeman 2006
Phil Evans 2005
Mariann Bienz 2003
Venki Ramakrishnan 2003
John Kilmartin 2002
Andrew Leslie 2001
Cyrus Chothia 2000
Michel Goedert 2000
Kiyoshi Nagai 2000
John Walker 1995
Tony Crowther 1993
Michael Neuberger 1993
Jonathan Hodgkin 1990
Greg Winter 1990
Andrew McLachlan 1989
Barbara Pearse 1988
Hugh Pelham 1988
Terry Rabbits 1987
John Sulston 1986
Mark Bretscher 1985
Alan Fersht 1983
Richard Henderson 1983
Peter Lawrence 1983
Nigel Unwin 1983
Uli Arndt 1982
Dan Brown 1982
John Finch 1982
John D Smith 1976
César Milstein 1975
David Blow 1972
Brian Hartley 1971
Aaron Klug 1969
Sydney Brenner 1965
Hugh Huxley 1960
John Kendrew 1960
Francis Crick 1959
Max Perutz 1954
Fred Sanger 1954


Copley Medal Winners
Richard Henderson2016 – Richard Henderson
In recognition of his fundamental and revolutionary contributions to the development of electron microscopy of biological materials, enabling their atomic structures to be deduced.
2012 – John Walker
For his ground-breaking work on bioenergetics, discovering the mechanism of ATP synthesis in the mitochondrion.
1997 – Hugh Huxley
In recognition of his pioneering work on the structure of muscle and on the molecular mechanisms of muscle contraction, providing solutions to one of the great problems in physiology.
1991 – Sydney Brenner
In recognition of his many contributions to molecular genetics and developmental biology, and his recent role in the Human Genome mapping project.
1989 – César Milstein
In recognition of his outstanding contributions to immunology, in particular to the discovery of monoclonal antibodies and to the understanding of the role of somatic mutations in the maturation of the immune response.
1985 – Aaron Klug
In recognition of his outstanding contributions to our understanding of complex biological structures and the methods used for determining them.
1979 – Max Perutz
In recognition of his distinguished contributions to molecular biology through his own studies of the structure and biological activity of haemoglobin and his leadership in the development of the subject.
1977 – Fred Sanger
In recognition of his distinguished work on the chemical structure of proteins and his studies on the sequences of nucleic acids.
1975 – Francis Crick
In recognition of his elucidation of the structure of DNA and his continuing contribution to molecular biology.
Royal Medal
2011 – Greg Winter
For his pioneering work in protein engineering and therapeutic monoclonal antibodies, and his contributions as an inventor and entrepreneur.
2008 – Alan Fersht
For his seminal work in protein engineering, which he has developed into a fundamental tool in enzyme analysis and the problem of protein folding.
1985 – John Gurdon
For his outstanding contributions to the techniques of nuclear transplantation and the use of the amphibian egg for investigations on replication, transcription and translation of genes.
1982 – César Milstein
In recognition of his fundamental contribution to understanding the structure and genetic control of immunoglobulins; his hybridoma technique for producing monoclonal antibodies has revolutionized the potential practical applications of immunology.
1977 – Hugh Huxley
In recognition of his distinguished research on the structure of muscle and on the molecular mechanisms of contraction.
1974 – Sydney Brenner
In recognition of his distinguished contributions to molecular biology concerning the nature of the genetic code and its expression during development.
1972 – Francis Crick
In recognition of his elucidation of the structure of DNA and his continuing contribution to molecular biology.
1971 – Max Perutz
In recognition of his pioneering work on the molecular biology and structure of proteins.
1969 – Fred Sanger
In recognition of his pioneering work on the sequence of amino acids in proteins and of nucleotides of ribonucleic acids.
1965 – John Kendrew
In recognition of his distinguished contributions to the complete structural analysis of a protein molecule (myoglobin), particularly the biological aspects of this study.
Darwin Medal
2014 – John Sutherland
In recognition of his novel and convincing work on prebiotic chemistry, in particular his solution to the central problem of nucleoside synthesis.
1996 – John Sulston
In recognition of his leadership in the study of genome analysis with the potential to have a profound impact on the whole of biology.
1994 – Peter Lawrence
In recognition of his analysis of pattern formation during insect segmentation, and of his contribution to understanding how genetic processes specify spatial information.
Davy Medal
1998 – Alan Fersht
In recognition for his pioneering work on the analysis of proteins by combining the methods and ideas of physical-organic chemistry with those of protein engineering thus illuminating such processes as enzymatic catalysis, protein folding, protein-protein interactions and those macromolecule interactions in general that are dominated by the chemistry of the noncovalent bond.
Gabor Medal
1991 – Alan Fersht
In recognition of his pioneering work in the use of protein engineering to study protein structure and enzyme function.

Prize Lectures

Croonian Lecture
2007 – Aaron Klug
Engineered zinc finger proteins (ZFPs) for the regulation of gene expression.
2000 – Nigel Unwin
The nicotinic acetylcholine receptor and the structural basis of synaptic transmission.
1999 – Hugh Pelham
Intracellular membrane traffic: getting proteins sorted.
1989 – César Milstein
Antibodies, a paradigm of the biology of molecular recognition.
1986 – Sydney Brenner
The molecular genetics of muscle in the nematode Caenorhabditis elegant.
1976 – John Gurdon
Egg cytoplasm and gene control in development.
1975 – Fred Sanger
Nucleotide sequences.
1970 – Hugh Huxley
The structural basis of muscular contraction.
1968 – Max Perutz
The haemoglobin molecule.
1966 – Francis Crick
The genetic code.
Francis Crick Lecture
Madan Babu Mohan2016 – Madan Babu Mohan In recognition of Madan’s major and widespread contributions to computational biology.
2012 – Sarah Teichmann In recognition of Sarah’s exceptional achievements in structural bioinformatics relating to decoding the principles of protein interactions.
2009 – Jason Chin Reprogramming the code of life.
Leeuwenhoek Lecture
2012 – Brad Amos
In recognition of Brad’s exceptional impact on the field of cell biology through his co-development of the laser scanning confocal microscope.
2006 – Tony Crowther
Microscopy goes cold: frozen viruses reveal their structural secrets.
1973 – Aaron Klug
The structure and assembly of regular viruses.
Royal Society GlaxoSmithKline Prize and Lecture
2003 – Michael Neuberger
In recognition of his work on resolving the molecular mechanism of somatic antibody diversification, a key feature of immune response, with consequences reaching far beyond immunology to DNA instability and cancer.
1980 – César Milstein
In recognition of his pioneering the production of monoclonal antibodies from hybrid cell lines and initiating their application worldwide in many fields of biology and medicine.
Blackett Memorial Lecture/Jagdish Chandra Bose Memorial Lecture
1996 – Aaron Klug
Protein designs for the regulation of gene expression.
Florey Lecture
1992 – Hugh Pelham
The secretion of protein by cells.
Medawar Lecture
1992 – Max Perutz
Species adaptation in a protein molecule.


Mullard Award
Royal Society Mullard Award Confocal Group1994 – John White, Brad Amos, Richard Durbin and Michael Fordham
In recognition of their development of the MRC-600 series laser-scanning confocal imaging system, an ingenious and innovative means of improving the clarity and definition of microscopes.